End of year biomass Dr. Steven C. Pennings University of Houston

Department of Biology and Biochemistry University of Houston Houston Texas 77204-5513 USA

scpennin@central.uh.edu https://uh.edu/nsm/biology-biochemistry/people/profiles/steven-pennings/ https://orcid.org/0000-0003-4757-7125 Georgia Coastal Ecosystems LTER Project

Dept. of Marine Sciences University of Georgia Athens Georgia 30602-3636 USA

gcelter@uga.edu https://gce-lter.marsci.uga.edu/ Dr. Steven C. Pennings University of Houston

Department of Biology and Biochemistry University of Houston Houston Texas 77204-5513 USA

scpennin@central.uh.edu https://uh.edu/nsm/biology-biochemistry/people/profiles/steven-pennings/ https://orcid.org/0000-0003-4757-7125 Principal investigator 2000-01-01 Dr. Kazimierz Wieski University of Houston

Department of Biology and Biochemistry University of Houston Houston Texas 77204-5513 USA

trudnoswietnie@gmail.com Post Doctoral Associate Daniel F. Saucedo University of Georgia Marine Institute

University of Georgia Marine Institute Sapelo Island Georgia 31327 USA

dsaucedo@uga.edu Research technician 2006-09-06 Jacob Shalack University of Georgia Marine Institute

University of Georgia Marine Institute Sapelo Island Georgia 31327 USA

shalack@uga.edu Research technician 2007-06-01 Kristen Anstead University of Georgia Marine Institute

University of Georgia Marine Institute Sapelo Island Georgia 31327 USA

kanstead@uga.edu Research technician 2008-11-01 Hongyu Guo University of Houston

Department of Biology and Biochemistry 369 Science and Research Bldg 2 University of Houston Houston Texas 77204-5001 USA

greatuniverse@hotmail.com Graduate research assistant 2006-09-01 2026

S. Pennings (UH) and his lab monitor plant biomass with the goal of testing the hypothesis that end-of-year biomass varies as a function of 1) freshwater discharge from the Altamaha (especially in low-marsh plots), 2) local rainfall (especially in high-marsh plots), and 3) average sea level. In 2000 we set up permanent plots at all 10 LTER monitoring sites. Plots were established at creek-bank and mid-marsh sites (8 plots/zone/site). Most sites are dominated by Spartina alterniflora, but some zones at some sites are dominated by Juncus roemerianus, Spartina cynosuroides, or Zizaniopsis miliacea. An additional zone (high marsh Juncus) was established at site 10 in 2005 to increase replication of sites with Juncus. Plants have been non-destructively monitored (stem counts, heights, flowering status) in October of every year from 2000 to the present. Stem samples were taken adjacent to plots, measured, dried and weighed, in order to generate allometric regression relationships between height and mass.

A preliminary analysis of data for S. alterniflora (means and SE averaged over sites dominated by S. alterniflora) from 2000-2006 shows that end of year biomass in creekbank plots varied more than two-fold among years (Figure 6). The two years with highest biomass (2003 and 2005) were years with high discharge from the Altamaha River during the spring months, which reduced water column salinities and provided superior growing conditions. End-of-year biomass in the mid-marsh plots varied slightly less than two-fold among years and did not vary in synchrony with biomass in the creekbank plots (in particular, midmarsh plots showed no peak in biomass in 2003, Figure 6). Variation in biomass in mid-marsh plots is likely driven by variation in sea level and local precipitation, rather than Altamaha River discharge. These plots are also proving useful in documenting spatial and temporal variation in disturbance from physical (floating dead plant material called wrack) and biotic sources. Wrack disturbance is high in creekbank plots at some sites and low in mid-marsh plots. Biotic damage from snail grazing and pig digging is high in some mid-marsh plots at some sites. (Disturbed plots were omitted from above analyses of annual variation in biomass).

These plots are also proving useful in documenting spatial and temporal variation in disturbance from physical (floating dead plant material called wrack) and biotic sources. Wrack disturbance is high in creekbank plots at some sites and low in mid-marsh plots. Biotic damage from snail grazing and pig digging is high in some mid-marsh plots at some sites. (Disturbed plots were omitted from above analyses of annual variation in biomass).

Area 2: Patterns within the Domain (more information) Question 2: How do the spatial and temporal patterns of biogeochemical processes, primary production, community dynamics, decomposition, and disturbance vary across the estuarine landscape, and how do they relate to environmental gradients? (more information)

biomass discharge rainfall sea level LTER NSF Altamaha River Georgia GCE Juncus disturbance Overall geographic extent of the research project -81.493498 -81.195661 31.552554 31.304063 2000-01-01

National Science Foundation grant numbers OCE-9982133 and OCE-0620959

estuary marsh complex Eulonia - Transitional salt marsh/upland forest site at the upper reach of the Sapelo River near Eulonia, Georgia. The main marsh area is to the north of the channel where the upland is controlled by DNR. Several small creeks lie within the study area. Residential development is increasing on the upland areas south of the channel. A hydrographic sonde is deployed within this site attached to a private dock to the south of the main channel near the HW-17 bridge. The sonde is accessed from a dock on a private residence off of highway 17. As of late 2017 the property is owned by Linda Hornby. She asks that we notify her by text about an hour before arriving to service the sondes. There is no reason to go to this property other than for sonde work. -81.428481 -81.410732 31.546750 31.533963 Hunt Camp - Barrier island/marsh site on western Sapelo Island. This site is located at the upper reaches of the Duplin River, and is within the Sapelo Island National Estuarine Research Reserve. Existing well fields border small marsh area to northwest, some wells have been installed to south end of hammock where marsh is more extensive and permanent plots are located. Two existing hydrographic sondes and weather stations within this site are operated by SINERR (Hunt Camp dock) and UGAMI (flume dock). -81.295506 -81.258908 31.497026 31.462320 Four Mile Island - Low salt marsh/estuary site encompassing the southwestern end of Four Mile Island and the Sapelo River near Sapelo Sound. The large island is totally marsh, with no upland, and no development. Nearby areas, like "Sutherland Bluff", are developing with golf courses. A hydrographic sonde is deployed within this site attached to channel marker 4 (under U.S. Coast Guard permit). Salinity regime is similar to seawater. -81.318961 -81.292844 31.552554 31.533604 North Sapelo - High salinity marsh/Sapelo Sound site. Few or no small creeks, but one large creek is easily accessed. Further upstream upland drainage is affected by culvert at Reynolds' duck pond. Upland is heavily forested. Hydrographic sonde is deployed adjacent to this site attached to a channel marker (under U.S. Coast Guard permit). A huge mud flat is present at the northern tip of Sapelo Island. -81.233249 -81.195661 31.546868 31.515188 Meridian - Inland marsh and tidal creek site near Meridian, Georgia. The primary marsh site is to the south of the dock in front of the Sapelo Island Visitor Center. Some small creeks and one large creek (Hudson Creek) are present. Upland is heavily forested. Marsh to the north of the dock can be used for additional studies if larger areas are required. Upland to the North is being developed for residential use, so future access is uncertain. The USGS super station site (weather station plus multiple-sensor sonde) is deployed off the ferry dock. Salinity is similar to seawater because there is little freshwater input from the upland. -81.370752 -81.357256 31.463533 31.448419 Folly River - Salt marsh/tidal creek site at the Folly River near Doboy Sound. No upland areas present. The north side of Doboy Sound immediately opposite is similar, but creeks are not as nice. No existing markers are close enough for sonde deployment, so we will initially use the existing sonde at Marsh Landing operated by UGAMI for surrogate hydrographic data. -81.354413 -81.327201 31.442538 31.417640 Dean Creek - Doboy Sound/salt marsh site at the southern end of Sapelo Island near Dean Creek and the Sapelo lighthouse. A few small creeks are present, but a large creek (Dean Creek) is easily accessed. Upland is composed of small hammocks and some constructed causeways, with sand dune complexes east of Dean Creek and extending to the beach. Upper end of the watershed is affected by a culvert at Beach Road and heavy student use of marsh immediately adjacent to culvert. A hydrographic sonde is deployed in Doboy Sound near Commodore Island approximately 1.5km from this site. GCE6 is also the focus of Sapelo Island Microbial Observatory research on microbial diversity and genomics (http://simo.marsci.uga.edu) -81.299522 -81.264378 31.394072 31.371003 Carrs Island - Freshwater marsh site along the northern bank of Hammersmith Creek at the south end of Carrs Island. Upland area is heavily forested. Northern 3/4 of the island was diked for rice agriculture, but the southern 1/4 is undisturbed. A hydrographic sonde is deployed in the Altamaha River near Hammersmith Creek, adjacent to this site. -81.493498 -81.475900 31.348774 31.334364 Alligator Creek - Mid-estuary/salt marsh site on the west side of Friday Cap Creek along the Altamaha River. This is a brackish marsh with salinities around 14 PSU during drought, less than 5 PSU normally. A hydrographic sonde is deployed adjacent to this site in the Altamaha River near Alligator Creek, attached to the US Coast Guard Daymarker 2 pilings. -81.426272 -81.410634 31.317808 31.304063 Rockdedundy Island - Lower estuary tidal creek and marsh site at Rockdedundy Island west of Wolf Island. A hydrographic sonde is deployed approximately 2km from this site in the Altamaha River near Rockdedundy Island, attached to the US Coast Guard Daymarker 201 pilings. -81.346730 -81.326495 31.358142 31.339173 2009 GCE NSF Annual Report

Figure 6. End-of-year biomass at GCE monitoring sites dominated by Spartina alterniflora, 2000-2006. Data are means and SE of creek bank (solid circles) and mid-marsh (open circles) sites. https://gce-lter.marsci.uga.edu/public/resources/projects/Fig6AnnualReport09.jpg Liu, W. and Pennings, S.C. 2021. Variation in synchrony of production among species, sites and intertidal zones in coastal marshes. Ecology. (DOI: 10.1002/ECY.3278) https://esajournals.onlinelibrary.wiley.com/doi/10.1002/ecy.3278 Liu, W. and Pennings, S.C. 2019. Self-thinning and size-dependent flowering of the grass Spartina alterniflora across space and time. Functional Ecology. 33:1830-1841. (DOI: 10.1111/1365-2435.13384) Li, S., Hopkinson, C.S., Schubauer-Berigan, J.P. and Pennings, S.C. 2018. Climate drivers of Zizaniopsis miliacea biomass in a Georgia, U.S.A. tidal fresh marsh. Limnology and Oceanography. 63:2266-2276. (DOI: 10.1002/lno.10937) Li, S. and Pennings, S.C. 2016. Disturbance in Georgia salt marshes: variation across space and time. Ecosphere. 7(10):e01487. (DOI: 10.1002/ecs2.1487) http://onlinelibrary.wiley.com/doi/10.1002/ecs2.1487/abstract Pennings, S.C. 2005. Presentation: Physical forcing and variation in salt marsh plant productivity at multiple time scales. Ecological Society of America 2005 Meeting - Ecology at multiple scales, August 7-12, 2005, Montreal, Canada. POR-GCEM-2604 - Soil salinity and organic content at GCE-LTER vegetation monitoring plots in October 2025 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=867 PLT-GCEM-2511c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2025 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=870 PLT-GCEM-2411a - Fall 2024 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=845 POR-GCEM-2504 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2024 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=853 POR-GCES-2106 - Long-term soil salinity and organic content from the Georgia Coastal Ecosystems LTER Project on Sapelo Island, Georgia https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=761 PLT-GCES-1609 - Long-term Plant Biomass Monitoring Data from the Georgia Coastal Ecosystems LTER Project on Sapelo Island, Georgia https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=590 PLT-GCES-1609c - Long-term Plant Biomass Monitoring Data from Altamaha River Plant Transition Sites near the Georgia Coastal Ecosystems LTER Project on Sapelo Island, Georgia https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=759 PLT-GCEM-2411b - Fall 2024 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=846 PLT-GCEM-2411c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2024 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=847 PLT-GCEM-2311a - Fall 2023 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=810 PLT-GCEM-2311b - Fall 2023 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=811 PLT-GCEM-2311c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2023 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=812 POR-GCEM-2401 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2023 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=816 PLT-GCEM-1610 - Monthly Vegetation and Invertebrate Population Monitoring near the Georgia Coastal Ecosystems LTER Flux Tower https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=604 POR-GCEM-2301 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2022 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=798 PLT-GCEM-2211a - Fall 2022 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=794 PLT-GCEM-2211b - Fall 2022 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=795 PLT-GCEM-2211c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2022 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=796 PLT-GCEM-2111a - Fall 2021 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=772 PLT-GCEM-2111b - Fall 2021 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=773 PLT-GCEM-2111c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2021 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=774 POR-GCEM-2201 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2021 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=775 PLT-GCEM-2011c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2020 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=735 PLT-GCEM-2011a - Fall 2020 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=736 PLT-GCEM-2011b - Fall 2020 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=756 POR-GCEM-2101 - Soil salinity and organic content at GCE-LTER vegetation monitoring plots in October 2020 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=764 PLT-GCEM-2011 - Plant allometry at GCE sampling sites 1-10 in October between 2002 and 2020. https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=755 PLT-GCEM-1911c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2019 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=734 PLT-GCEM-1811c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2018 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=685 PLT-GCEM-1811a - Fall 2018 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=686 PLT-GCEM-1811b - Fall 2018 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=687 PLT-GCEM-1911a - Fall 2019 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=704 PLT-GCEM-1911b - Fall 2019 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=705 PLT-GCEM-1711a - Fall 2017 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=639 PLT-GCEM-1711b - Fall 2017 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=640 PLT-GCEM-1711c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2017 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=636 POR-GCEM-1701 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2016 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=706 PLT-GCEM-1611a - Fall 2016 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=632 PLT-GCEM-1611c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2016 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=634 PLT-GCEM-1611b - Fall 2016 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=635 POR-GCEM-1801 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2017 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=707 POR-GCEM-1901 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2018 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=708 POR-GCEM-2001 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2019 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=709 POR-GCEM-1601 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2015 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=579 PLT-GCEM-1511a - Fall 2015 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=577 PLT-GCEM-1511b - Fall 2015 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=578 PLT-GCEM-1511c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2015 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=576 PLT-GCEM-1412a - Fall 2014 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=472 PLT-GCEM-1412b - Fall 2014 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=473 PLT-GCEM-1412c - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2014 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=479 POR-GCEM-1312 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2013 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=487 POR-GCEM-1411 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2014 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=488 PLT-GCEM-1403a - Fall 2013 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=456 PLT-GCEM-1403b - Fall 2013 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=457 PLT-GCEM-1312 - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2013 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=478 PLT-GCEM-1311a - Fall 2012 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=454 PLT-GCEM-1311b - Fall 2012 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=455 PLT-GCEM-1210 - GCE-LTER Altamaha River Plant Community Monitoring Survey in October 2012 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=431 POR-GCEM-1212 - Soil salinity at GCE-LTER vegetation monitoring plots in October 2012 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=433 POR-GCEM-1207 - Soil salinity and organic content at GCE-LTER vegetation monitoring plots in October 2011 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=425 PLT-GCEM-1112a - Fall 2011 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=336 PLT-GCEM-1112b - Fall 2011 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=337 PLT-GCEM-1010a - Fall 2010 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=325 PLT-GCEM-1010b - Fall 2010 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=326 POR-GCEM-1011 - Soil salinity and water content at GCE-LTER vegetation monitoring plots in October 2010 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=328 POR-GCEM-1006 - Soil salinity and organic content at GCE-LTER vegetation monitoring plots in October 2009 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=327 PLT-GCEM-1004a - Fall 2009 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=319 PLT-GCEM-1004b - Fall 2009 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=320 PLT-GCEM-0812a - Fall 2008 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=312 PLT-GCEM-0812b - Fall 2008 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=313 PLT-GCEM-0801a - Fall 2007 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=303 PLT-GCEM-0801b - Fall 2007 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=304 PLT-GCEM-0612a - Fall 2006 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=280 PLT-GCEM-0612b - Fall 2006 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=281 PLT-GCEM-0511a - Fall 2005 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=266 PLT-GCEM-0511b - Fall 2005 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=267 PLT-GCEM-0501a - Fall 2004 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=230 PLT-GCEM-0501b - Fall 2004 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=231 PLT-GCEM-0311a - Fall 2003 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=178 PLT-GCEM-0311b - Fall 2003 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=179 PLT-GCEM-0211 - Fall 2002 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=26 PLT-GCEM-0303c - Fall 2002 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=94 PLT-GCEM-0110 - Fall 2001 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=8 PLT-GCEM-0303b - Fall 2001 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=93 PLT-GCEM-0303a - Fall 2000 plant monitoring survey -- biomass calculated from shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=92 PLT-GCEM-0101 - Fall 2000 plant monitoring survey -- shoot height and flowering status of plants in permanent plots at GCE sampling sites 1-10 https://gce-lter.marsci.uga.edu/public/app/dataset_details.asp?id=2 Figure 6. End-of-year biomass at GCE monitoring sites dominated by Spartina alterniflora, 2000-2006. Data are means and SE of creek bank (solid circles) and mid-marsh (open circles) sites. https://gce-lter.marsci.uga.edu/public/resources/projects/plantBiomassQ2.png