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<shortName>Rainfall variation and marsh plants</shortName>
<title>Effect of rainfall variation on high marsh plants</title>
<creator>
<individualName>
<salutation>Dr.</salutation>
<givenName>Steven</givenName>
<givenName>C.</givenName>
<surName>Pennings</surName>
</individualName>
<organizationName>University of Houston</organizationName>
<address>
<deliveryPoint>Department of Biology and Biochemistry</deliveryPoint>
<deliveryPoint>University of Houston</deliveryPoint>
<city>Houston</city>
<administrativeArea>Texas</administrativeArea>
<postalCode>77204-5513</postalCode>
<country>USA</country>
</address>
<electronicMailAddress>scpennin@central.uh.edu</electronicMailAddress>
<onlineUrl>https://uh.edu/nsm/biology-biochemistry/people/profiles/steven-pennings/</onlineUrl>
<userId directory="https://orcid.org">https://orcid.org/0000-0003-4757-7125</userId>
</creator>
<metadataProvider><organizationName>Georgia Coastal Ecosystems LTER Project</organizationName>
<address>
<deliveryPoint>Dept. of Marine Sciences</deliveryPoint>
<deliveryPoint>University of Georgia</deliveryPoint>
<city>Athens</city>
<administrativeArea>Georgia</administrativeArea>
<postalCode>30602-3636</postalCode>
<country>USA</country>
</address>
<electronicMailAddress>gcelter@uga.edu</electronicMailAddress>
<onlineUrl>https://gce-lter.marsci.uga.edu/</onlineUrl>
</metadataProvider>
<associatedParty>
<individualName>
<salutation>Dr.</salutation>
<givenName>Steven</givenName>
<givenName>C.</givenName>
<surName>Pennings</surName>
</individualName>
<organizationName>University of Houston</organizationName>
<address>
<deliveryPoint>Department of Biology and Biochemistry</deliveryPoint>
<deliveryPoint>University of Houston</deliveryPoint>
<city>Houston</city>
<administrativeArea>Texas</administrativeArea>
<postalCode>77204-5513</postalCode>
<country>USA</country>
</address>
<electronicMailAddress>scpennin@central.uh.edu</electronicMailAddress>
<onlineUrl>https://uh.edu/nsm/biology-biochemistry/people/profiles/steven-pennings/</onlineUrl>
<userId directory="https://orcid.org">https://orcid.org/0000-0003-4757-7125</userId>
<role>Principal investigator</role>
</associatedParty>
<pubDate>2026</pubDate>
<abstract>
<section>
<title>Overview</title>
<para>Pennings (UH)  is testing the hypothesis that annual variation in high-marsh plant species composition is driven by variation in rainfall.  Salt marsh vegetation often consists of discrete stands with abrupt borders between different species or associations.  Pennings has monitored mid-summer plant composition at permanent plots located on 3 types of vegetation borders (Spartina alterniflora-Juncus roemerianus, S. alterniflora-meadow, meadow-Juncus roemerianus), at two sites each, since 1996.  Vegetation composition in these plots is dynamic and appears to be related to variation in rainfall, although again more years of data will be needed to test this hypothesis rigorously.</para>
</section>
<section>
<title>Associated GCE LTER research questions</title>
<para>Area 2: Patterns within the Domain (<ulink url="http://gce-lter.marsci.uga.edu/public/research/gce3_area2.asp">more information</ulink>)</para>
<para>Question 2: How do the spatial and temporal patterns of biogeochemical processes, primary production, community dynamics, decomposition, and disturbance vary across the estuarine landscape, and how do they relate to environmental gradients? (<ulink url="http://gce-lter.marsci.uga.edu/public/research/gce2_q2.asp">more information</ulink>)</para>
</section>
</abstract>
<keywordSet name="measurement">
<keyword>rainfall</keyword>
</keywordSet>
<keywordSet name="organization">
<keyword>LTER</keyword>
<keyword>NSF</keyword>
</keywordSet>
<keywordSet name="place">
<keyword>Georiga</keyword>
</keywordSet>
<keywordSet name="site">
<keyword>GCE</keyword>
</keywordSet>
<keywordSet name="taxonomic">
<keyword>Juncus</keyword>
<keyword>Spartina</keyword>
</keywordSet>
<keywordSet name="theme">
<keyword>salt marsh</keyword>
<keyword>vegetation</keyword>
</keywordSet>
<coverage>
<geographicCoverage>
<geographicDescription>Overall geographic extent of the research project</geographicDescription>
<boundingCoordinates>
<westBoundingCoordinate>-81.297445</westBoundingCoordinate>
<eastBoundingCoordinate>-81.289170</eastBoundingCoordinate>
<northBoundingCoordinate>31.418899</northBoundingCoordinate>
<southBoundingCoordinate>31.400830</southBoundingCoordinate>
</boundingCoordinates>
</geographicCoverage>
<temporalCoverage>
<ongoing>
<beginDate>
<calendarDate>1996-01-01</calendarDate>
</beginDate>
</ongoing>
</temporalCoverage>
</coverage>
<funding>
<section>
<para>National Science Foundation grant numbers OCE-9982133 and OCE-0620959</para>
</section>
</funding>
<studyAreaDescription>
<descriptor name="hydrology" citableClassificationSystem="false">
<descriptorValue>estuary marsh complex</descriptorValue>
</descriptor>
<coverage>
<geographicCoverage>
<geographicDescription>Marsh Landing - Marsh landing, where the Georgia DNR ferry docks on its route between the mainland (Meridian) and Sapelo Island.</geographicDescription>
<boundingCoordinates>
<westBoundingCoordinate>-81.297445</westBoundingCoordinate>
<eastBoundingCoordinate>-81.294606</eastBoundingCoordinate>
<northBoundingCoordinate>31.418899</northBoundingCoordinate>
<southBoundingCoordinate>31.416399</southBoundingCoordinate>
</boundingCoordinates>
</geographicCoverage>
<geographicCoverage>
<geographicDescription>Shell_Hammock - Shell_Hammock sampling site used by Amy Kunza in 2005 to study plant species richness</geographicDescription>
<boundingCoordinates>
<westBoundingCoordinate>-81.289170</westBoundingCoordinate>
<eastBoundingCoordinate>-81.289170</eastBoundingCoordinate>
<northBoundingCoordinate>31.400830</northBoundingCoordinate>
<southBoundingCoordinate>31.400830</southBoundingCoordinate>
</boundingCoordinates>
</geographicCoverage>
</coverage>
</studyAreaDescription>
<associatedMaterial category="publication" id="gce.44" date="2001">
<distribution>
<offline>
<mediumName>Pennings, S.C., Bertness, M.D., Donnelly, J.P., Ewanchuk, P.J., Silliman, B.R. and Callaway, R.M. 2001. Presentation: Impacts of global change on coastal salt marshes. Keynote address to the German Limnological Association, September 17-21, 2001, Kiel, Germany.</mediumName>
</offline>
</distribution>
</associatedMaterial>
</lter:researchProject>
